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Nevertheless, two major phylogeographic groupings were discovered in dholes of the Asian mainland, which likely diverged during a glaciation event.One population extends from South, Central, and North India (south of the Ganges) into Burma, and the other extends from India north of the Ganges into northeastern India, Burma, Thailand and the Malaysian Peninsula.The throat, chest, flanks, and belly and the upper parts of the limbs are less brightly coloured, and are more yellowish in tone.

It resembled Canis in its physical form more than the modern species, which has greatly reduced molars, whose cusps have developed into sharply trenchant points. The dhole became extinct in much of Europe during the late Würm period, The dhole's distinctive morphology has been a source of much confusion in determining the species' systematic position among the Canidae.The origin of dholes in Sumatra and Java is, as of 2005, unclear, as they show greater relatedness to dholes in India, Burma and China rather than with those in nearby Malaysia.In the absence of further data, the researchers involved in the study speculated that Javan and Sumatran dholes could have been introduced to the islands by humans.Playful dholes open their mouths with their lips retracted and their tails held in a vertical position whilst assuming a play bow.Aggressive or threatening dholes pucker their lips forward in a snarl and raise the hairs on their backs, as well as keep their tails horizontal or vertical.

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The amphibians evolved in the Devonian period (about 370 million years ago) as the first vertebrates to occupy the land.

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